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Apistogramma Species List

By Species-Groups/Complexes



As of November 2018






Compiled by Mike Wise
(with assistance from Frank Hättich
and Tom Christoffersen)



The genus Apistogramma is the most species-rich genus of cichlid fishes in South America. When I first compiled a list of Apistogramma species for the Apistogramma Study Group back in 1985 there were only 41 scientifically described species, one of which is now considered a junior synonym, and less than 10 species that we considered undescribed. Things have become more complicated since then.

Presently, Fishbase lists 90 scientifically described species as valid, but there are many more undescribed Apistogramma species that have entered the hobby over the past 35 years. In addition, a large number of distinct populations and even some color morphs (both often called "forms") exist that may or may not be species in their own right. At the time of this writing we have recognized almost 440 species, forms, and distinct populations of Apistogramma.

The following list is the compilers’ opinion of which Apistogramma forms are distinct and separate species, and which are merely geographic populations. Our list is based on information provided to us by ichthyologists and apistophiles from all over the world. It was originally influenced strongly by the works of Kullander, Koslowski and preliminary genetic work by Miller and Schliewen. New data produced under the auspices of Institute of Research for Development – Instituto de Investigaciones de la Amazonía Peruana (IRD-IIAP) by Römer, Tougard and other ichtyologists have expanded our knowledge even more.

I think that there needs to be some explanations for first-time users.

Why are the species listed by species-groups instead of alphabetically?
The primary purpose of this list is to help identify your apistos. With so many species/forms, I decided long ago that the list would be more useful for identification purposes if all similar species were listed together. I adhere to the "divide and conquer" method of identification. Why wade through almost 440 species/forms in an alphabetical list if I can only compare similar species/forms? In that way there are many fewer apistos that you need to consider. To help you understand species-groups I describe the various species-groups in an other article on this website.

Can there really be so many species?
The compilers of this list would like to think that there are fewer, but recent published and unpublished genetic data that we have indicates that there actually are as many – if not more – species than we show on the list. Therefore we have resigned ourselves to be "splitters". We do this for two reasons. First, genetic studies indicate that the genus Apistogramma is not only evolving very rapidly (Farias, et al., 2001), but that genetically distinct species can be almost indistinguishable from each other (Ready, et al., 2006). Second, we feel that we need to avoid crossing different populations in the hobby as much as possible. Many of these populations are extremely rare and have not yet been studied. Once crossed with other similar populations many of these possible species could be lost to the hobby. If they turn out to be populations of the same species then no real harm is done, but if they turn out to be distinct species we then lose species purity in the hobby due to unintentional hybridization.

How to read the list?
Valid scientifically described species are listed in alphabetical order first in each species-group, complex or subcomplex. Apistogramma "forms" that we believe are distinct species, but scientifically undescribed are then listed in alphabetical order. Forms that we believe are only geographic populations of a species are indicated as such by being listed below the species to which we believe they belong, but they are shown indented. Many species have been given more than one name. These are listed in "Other Names". Where there is more than one common name for an undescribed species or form, we tend to use the oldest or most popular published name, as opposed to commercial/trade names. There are exceptions, however. We try to avoid using names that look like scientifically described names, although they are not (e.g. Albertini, Wilhelmi). Such common names continue to cause confusion among some hobbyists about what is a valid scientifically described species and what is not. Instead, we give preference to location names, followed by descriptive names. We also try to avoid common names that might be confusing, such as our using the name A. sp. "Cruzeiro" (for the city of Cruzeiro do Sul) instead of A. sp. "Juruá", which can be confused with A. juruensis. A-numbers listed in Stawikowski (2005) are included for most species. Those without A-numbers have been introduced into the hobby after its publication. D-numbers are used for species/forms that have been discovered and cataloged by Daniel Mejia in cooperation with Tom Christoffersen, many now appearing in the hobby trade. Where a name is listed as = A. xxx? it indicates that we are not completely sure if it is the same species. Finally, we provide a generalized "Distribution" for each form, if known.

We doubt that others with an extensive knowledge of the genus will agree completely with this list, but we feel that it is accurate for the most part. It will undoubtedly change – as it has in the past – as new species and data appear. We – Tom, Frank and I – hope that the list is helpful to those using it. We are always interested in others’ opinions. Such discussions can only improve our understanding of the many species of Apistogramma.

– Mike Wise      






Phylogeny of the Genus Apistogramma





   A.-regani-Lineage
      A.-regani-Group
          A.-caetei -Complex
          A.-sp.-"Xingu"-Complex
          A.-kullanderi-Complex
          A.-resticulosa-Complex
          A.-commbrae-Complex
          A.-borellii-Complex
          Apistogrammoides
          A.-regani-Complex
          A.-sp.-"Angle-patch"-Complex
          A.-eunotus-Complex

                A.-eunotus-Subcomplex
                A.-cruzi-Subcomplex
                A.-ortegai-Subcomplex

     A.-alacrina-Group
     A.-macmasteri-Group

          A.-hongsloi-Complex
          A.-macmasteri-Complex

                A.-macmasteri-Subcomplex
                A.-hoignei-Subcomplex


   A.-pertensis-Lineage
         A. wapisana
         A.-steindachneri-Group
         A.-pertensis-Group

              A.-pertensis-Complex
              A.-velifera-Complex
         A.-iniridae-Group


   A.-trifasciata-Lineage
      A.-trifasciata-Sublineage
       Aarua
       A.-brevis-Group
       A.-sp.-"D10"-Group
       A.-nijsseni-Group
       A.-cacatuoides-Group
       A.-norberti-Group
       A.-atahualpa-Group
            A.-atahualpa-Complex
            A.-barlowi-Complex

       A.-trifasciata-Group
      A.-agassizii-Sublineage
       A.-elizabethae-Group
       A.-gibbiceps-Group
       A.-bitaeniata-Group
            A.-bitaeniata-Complex
            A.-paucisquamis-Complex

       A.-agassizii-Group
            A.-pulchra-Complex
            A.-agassizii-Complex


  A.-diplotaenia-Lineage
        A.-diplotaenia-Group


       Taeniacara
















































A.-regani-Lineage

A.-regani-Group

A.-caetei-Complex

    Name     Other Names A-No.     Distribution
A. caetei

   A. cf. caetei (Gurupí)

   A. cf. caetei (Turiaçu)


= A. caetei?

= A. caetei?
   - 

A 26

A 29
Rio Caeté; Rio Apeú (N. E. Coast Brazil)

Rio Gurupi (Brazil)

Rio Turiaçu (Brazil)
A. cf. caetei (Guamá)

   A. cf. caetei (Capim)
A 27

A 28
Rio Guamá (L. Rio Capim, Brazil)

Rio Capim (Brazil)
A. cf. caetei (Maramanim)    -  Rio Marapanim (Brazil)
A. cf. caetei (Paraguay) A. sp. "Paraguay";
A. sp. "Paraguay I"
A 31 Rio Paraguay? (Brazil?, Paraguay?)
A. cf. caetei (Rotwangen/Red-cheeks) A. cf. caetei Marajó A 30 Ilha de Marajó; Rio Capim?; Rio Turiaçu? (Brazil)
A. sp. "Araguaia"



   A. sp. "Araguaia" (Crixas)

   A. sp. "Araguaia" (Marabá)
A. sp. "Aruana";
A. sp. "Araguaia" (Aruanã);
A. sp. "Araguaia" (Santana)

A. sp. "Crixas"
A 22



A 23

A 24
L. Tocantins; Rio Araguaia (Aruanã, Brazil)



Rio Crixas-Mirim (U. Araguaia, Brazil)

Rio Araguaia near Marabá (Brazil)
A. sp. "Macapá" A 32 Amazon delta (Near Macapá, Brazil)



A.-regani-Group

A.-sp.-"Xingu" / A.-caetei-Complex Bridge?

    Name     Other Names A-No.     Distribution
A. piauiensis A. sp. "Paraguay III" A 21 Rio Parnaiba (N.E. Coast, Brazil)
A. rubrolineata (Bolivia)

   A. rubrolineata (Peru)
A. sp. "Beni"

A. sp. "Manu"
A 18

A 19
Río Beni system (Bolivia)

Río Manuripe (Peru)
A. tucurui A. sp. "Tucurui" A 20 L. Rio Tocantins (Tucurui, Brazil)


A.-regani-Group

A.-sp.-"Xingu"-Complex

    Name     Other Names A-No.     Distribution
A. sp. "Cajueiro"    - L. R. Teles Pires near Cajueiro ferry; R. Paranaita; R. São Benedito (Brazil)
A. sp. "Chao"


   A. sp. "Itaparaná"
A. sp. "Rio Xingu"
(Chao 93-107)

= A. sp. "Chao"?;
A. sp. "Kinnfleck/Chin-spot"
A 42


A 43
M. Rio Xingu (S of Altamira, Brazil)


Reported to be from the Rio Itaparaná
(M. Rio Purus), but probably from the
Rio Xingu (Brazil)
A. sp. "Jabuti" A. taeniata Santarém A 36 Rio Jabuti (near Santarém, Brazil)
A. sp. "Jamanxim"    - Rio Jamanxim (U. Rio Xingu, Brazil)
A. sp. "Middle Curuá"    - L. R.Treze de Maio (U. Rio Xingu, Brazil)
A. sp. "Parati" A. sp. "Xingu III"

A. sp. "Paje"
A 40

A 41
L. Rio Xingu (Vitoria do Xingu to Altamira, Brazil)
A. sp. "Xingu"



   A. cf. sp. "Xingu" (Gold Nugget)

   A. sp. aff. "Peixoto"


   A. sp. "Vielfleck/Multi-spot"
A. sp. "Rio Xingu" [Stawikowski];
A. sp. "Xingu" (Red-lobes)

 

A. sp. aff. "Peixoto" (Purus)


A. sp. "Amapa Guttata";
A. sp. "Brazil-Guttata";
A. sp. "Pineapple";
A. sp. "Samaúma"
A 37



A 37a

   - 


A 38
L. Rio Xingu (near Altamira, Brazil)



L. Rio Xingu (Brazil)

Reportedly from the Rio Purus,
but probably from the Rio Xingu (Brazil)

L. Rio Xingu (Altamira)
& Rio Tucurui do Xingu (Vitoria, Brazil)


A.-regani-Group

A.-kullanderi-Complex

    Name     Other Names A-No.     Distribution
A. eleutheria A. sp. "Upper Treze de Maio"    - Rio Treze de Maio
(U. Rio Xingu, Brazil)
A. flavipedunculata A. sp. "Upper Curuá"    - Rio Treze de Maio (U. Rio Curuá, Brazil)
A. kullanderi A. sp. "Gigas"    - U. Rio Curuá (U. Rio Xingu, Brazil)
A. sp. "Peixoto" A. sp. "Rio Peixoto-Azevedo";
A. sp. "Serpa";
= A. sp. "Teles Pires"?
A 39 Rio Peixoto de Azevedo (U. Rio São Manuel ou Teles Pires, Brazil)
A. sp. "Teles Pires" = A. sp. "Peixoto"?    - M. Rio São Manuel ou Teles Pires (Brazil)


A.-regani-Group

A.-resticulosa-Complex

    Name     Other Names A-No.     Distribution
A. resticulosa

    A. cf. resticulosa (Nathaneal)
A 12

A 14
U. Rio Madeira (Humaitá, Brazil)

Rio Maçangana (Near Alto Paraiso, Brazil)
A. cf. resticulosa (Aripuanã) A 15 Rio Aripuanã near Novo Aripuanã
(M. Rio Madeira, Brazil)
A. cf. resticulosa (Humaitá) A 13 U. Rio Madeira (Humaitá, Brazil)
A. cf. resticulosa (Lagoa Abunã) A. sp. "Lagua Abunã"    - Lagoa Abunã (M. Rio Abunã,
Plácido de Castro, Brazil)
A. cf. resticulosa (Madre de Dios)    -  U. R. Madre de Dios (Bolivia/Peru)
A. cf. resticulosa (Madeira) A 17 L. Rio Madeira (Brazil)
A. cf. resticulosa (Mamoré Blue)



   A. cf. resticulosa (Guaporé 1)
= A. sp. "Malome"?;
A. sp. aff. taeniata
(Römer CA2)

A. sp. "Guaporé 1"
A 16



   -
L. Rio Mamoré (Brazil/Bolivia)



L.-M. Rio Guaporé near Vila Bela (Brazil)
A. cf. resticulosa (Rio Beni)    - L. Rio Beni near Riberalta (Bolivia)
A. taeniata

   A. cf. taeniata 1 (Curuá)

   A. cf. taeniata 1 (Curuá-Una)

   A. cf. taeniata 1 (Prainha)

   A. cf. taeniata 1 (Tapajós)

   A. cf. taeniata 1 (Xingu)




= A. sp. "Rio Iriri"?
A 1

A 7

A 6

A 8

A 2

A 9
Rio Cupari (L. Rio Tapajós,Brazil)

Rio Curuá (Brazil)

L. Rio Curuá-Una (Brazil)

Prainha (Brazil)

L. Rio Tapajós (Brazil)

L. Rio Xingu (Brazil)
A. cf. taeniata 2 (Igarapé Açu)


   A. cf. taeniata 2 (Arapiuns)

   A. cf. taeniata 2
      (Blauglanz/Blue-sheen)

   A. cf. taeniata 2 (Mayland & Bork)


   A. cf. taeniata 2 (Óbidos)
= A. cf. taeniata (Alenquer)?
= A. sp. "Alenquer"?



A. sp. "Blauglanz/Blue-sheen"


= A. cf. taeniata (Igarapé Açu)
   w/immaculate caudal?
A 4


A 3

A 10


A 4?


A 5
M. Amazon between Manaus & Santarém
(Brazil)

Rio Arapiuns (L. Rio Tapajós, Brazil)

Middle Amazon (Brazil)


L. Amazon (Brazil)


Near Óbido (Brazil)
A. sp. "Wangenflecken/Cheek-spots" A 11 Ilha de Marajó (Brazil)


A.-regani-Group

A.-resticulosa-Complex?

    Name     Other Names A-No.     Distribution
A. pleurotaenia A 33 Río Paraguay (Paraguay? Argentina?)
A. urteagai A 35 Río Madre de Dios (SE Peru & SW Brazil)
A. sp. "Mosaik/Mosaic" A 34 Peru import; no precise location
A. sp. "Kosho-Kosho" A. sp. "Puerto Maldonado"    - Río Madre de Dios
(East of Puerto Maldonado, Peru)


A.-regani-Group

A.-resticulosa / caetei-Complex?

    Name     Other Names A-No.     Distribution
A. sp. "Paru" A 25 Rio Paru above the Cachoeira Parnagua (Brazil)
A. sp. "Steel-blue" A. sp. "Blaukopf/Blue-head";
A. sp. "New Blue"
   - resticulosa X caetei - Domestic Hybrid?


A.-regani-Group

A.-commbrae-Complex

    Name     Other Names A-No.     Distribution
A. commbrae

   A. commbrae
      (w/separate caudal spot)

   A. cf. commbrae (Salto)
A 93

A 94


   -
U. & M. Río Paraguay (Paraguay & Brazil)

Import from Paraguay


Río Uruguay (near Salto, Uruguay)
A. inconspicua (Bolivia)


   A. cf. inconspicua (Brazil)

   A. sp. aff. similis





= A. inconspicua?
A 96


A 97

   - 
U. Rio Paraguai (Brazil);
U. Río Guaporé (Brazil/Bolivia)

U. Rio Guaporé (Brazil)

Río Itonamas & San Martín (Bolivia)
A. linkei (Santa Cruz)

   A. cf. linkei (Cochabamba)

   A. linkei (Río Itonamas)
A. sp. "Gelbbrust/Yellow-breast" A 99

   - 

A 98
Río Mamoré & Río San Miguel (Bolivia)

Río Chaparé near Cochabamba (Bolivia)

Río Itonamas (Guaporé system) (Bolivia)
A. similis A. sp. "Goldlippe/Gold-lips" A 95 Laguna La Brava, Río Yata system (Bolivia)
A. sp. "Mariae"    -  commbrae x ? Hybrid Domestic Form


A.-regani-Group

A.-borellii-Complex

    Name     Other Names A-No.     Distribution
A. borellii




   A. cf. borellii (Bolivia)

   A. cf. borellii (Opal/Opalita)


   A. cf. borellii (Paraguay)

   A. cf. borellii (Río Uruguai)
A. aequipinnis; A. reitzigi;
A. ritense; A. rondoni;
A. sp. "Opal/Opalita";
= A. sp. "Paraguay II"?
A 101




A 102

A 105


A 103

A 104
M.-U. Rio Paraguay (Brazil, Paraguay,
N. Argentina); Mato Groso (Brazil, Bolivia)



Mato Groso (Brazil, Bolivia)

Río Paraguay (Brazil, Paraguay,
N. Argentina); Mato Groso (Brazil, Bolivia)

M.-U. Río Paraguay (Paraguay)

M.-U. Río Uruguai (Brazil, Uruguay)
A. sp. "Paraguay II" = A. borellii?    -  L. Río Paraguay? (N. Argentina)?


A.-regani-Group

Apistogrammoides

    Name     Other Names A-No.     Distribution
Apistogrammoides pucallpaensis A 100 M. - L. Río Ucayali (Peru) to U. Río Amazonas (Peru & Colombia)


A.-regani-Group

A.-regani-Complex

    Name     Other Names A-No.     Distribution
A. acrensis A. sp. "Rio Branco";
A. sp. "Rio Acre"
A 63 Rio Acre (near Rio Branco,Brazil)
A. geisleri

   A. geisleri (Parintins)

   A. geisleri (Smaragd/Emerald)
A 58

A 60

A 59
Rio Curuçambá & L. Rio Tapajós (Brazil)

Rio Parintins (L. Amazon,Brazil)

L. Amazon (NW of Santarém,Brazil)
A. cf. geisleri (Brazil) A. sp. "Brazil" A 61 L. Amazon?
A. ortmanni

   A. cf. ortmanni (aquarium import)

   A. cf. ortmanni (rupununi River)
A 46

A 48

A 47
Guyana & Western Surinam

Aquarium import (Guyana)

Rupununi River (Guyana
A. cf. ortmanni (Río Cuyuni) A 49 Cuyuni River & Tumeremo Reservoir
(Eastern Venezuela)
A. paulmuelleri A. sp. "Masken/Masked";
A. sp. "Apache";
A. sp. "Carapintada" (in part);
A. sp. "Peru (II)";
A. cf. regani (Belem)
A 52 L. Río Ucayali (Contamaná, Peru)
A. cf. paulmuelleri (Juruá) A. sp. "Eirunepé";
A. cf. "Masken" (Juruá)
   -  M. Rio Juruá (Eirunepé, Brazil)
A. cf. paulmuelleri (Peru-Regani) Large A. sp. "Peru-Regani" (= less
colorful population of
A. paulmuelleri?)
   -  U. Río Tigre (Peru)
A. cf. paulmuelleri (Peru-Regani) Small    -  M. Río Ucayali (Peru)
A. regani A 55 L. Rio Negro; M. Amazon (Brazil)
A. cf. regani (Trombetas) A 56 Igarapé São Miguel, Rio Trombetas (Brazil)
A. sp. "Barcelos" A. sp. "Solimões" A 57 M. Rio Negro (Barcelos, Brazil),
L. Solimões (Brazil)
A. sp. "Cuiari" A. sp. "Kuiari"    -  R. Cuiari (Colombia, Brazil?)
A. sp. "Gelbwangen/Yellow-cheeks"


   A. cf. sp. "Gelbwangen/
      Yellow-cheeks"
      (Sichelbinden/Sickle-bands)
A. sp. "Rio Solimões";
A. sp. "Prahina"

A. sp. "Sichelbinden/
Sickle-bands"
(Aqualog Extra)
A 62


   - 
M. – L. Amazon (between Manaus & Prainha, Brazil)

Unknown, M. – L. Amazon (Brazil)?
A. sp. "Rotporen/Red-pores" A. sp. "Amapá Bitter I";
"CA 96 Amapa"
A 54 Near Lourenço, Amapá, Brazil
A. sp. "São Gabriel" A. sp. "Alto Negro";
= A. sp. "Tiquié regani"?
A 53 U. Rio Negro near São Gabriel da Cachoeira (Brazil)


A.-regani-Group

A.-sp.-"Winkelfleck/Angle-patch"-Complex

    Name     Other Names A-No.     Distribution
A. sp. "Winkelfleck/Angle-spot 1" A. sp. "Carapintada" (in part);
A. sp. "Winkelfleck" (light);
A. sp. "Tiger Stripes"
A 44 L. Río Marañón (Nauta, Peru)
A. sp. "Winkelfleck/Angle-spot 2" A. sp. "Oroja";
A. sp. "Winkelfleck" (dark)
A 45 Río Orosa (Peru)


A.-regani-Group

A.-regani-? / A.-eunotus-Complex?

    Name     Other Names A-No.     Distribution
A. gossei


   A. cf. gossei
A 50


A 51
Río Oyapock/Oiapoque
(Brazil/French Guiana)

French Guiana
A. moae A 72 Rio Moa (U. Rio Juruá, Brazil)
A. sp. "Amapá" (Rio Amapá Grande)


   A. sp. "Amapá" (Fortaleza)


   A. sp. "Amapá" (Rio Araguari)

   A. sp. "Amapá" (Rio Caçiporé)

   A. sp. "Amapá" (Rio Calçoene)

   A. sp. "Amapá" (Rio Macari)



A. sp. "Fortaleza"
(Aqualog Extra)
A 68


   - 


A 67

A 71

A 70

A 69
Rio Amapá Grande,
Amapá State(Brazil)

Near Fortaleza, Amapá (Brazil)


Rio Araguari, Amapá State (Brazil)

Rio Caçiporé, Amapá State (Brazil)

Rio Calçoene, Amapá State (Brazil)

Rio Macari, Amapá State (Brazil)
A. sp. "Rio Candeias" A 64 Rio Candeias (U. Rio Madeira, Brazil)
A. sp. "Rio Jaru" A 65 Rio Jaru (U. Rio Madeira, Brazil)
A. sp. "Xipamanu" A 66 Rio Xipamanu/Chipamanu
(U. Rio Abunã, Brazil)


A.-regani-Group

A.-eunotus-Complex

A.-eunotus-Subcomplex

    Name     Other Names A-No.     Distribution
A. cinilabra





   A. (cf.) cinilabra
      (with no caudal pattern)
A. sp. "Schwarzbrust/
Black-breast";
A. sp. "2000-1"(panduro);
A. sp. "Roterpunkt/Redder-spot"; A. sp. "Doña Martina"

A. sp. "KM 77"
A 80





   -
Río Itaya drainage near Nauta (Peru)





KM 77 marker on Iquitos-Nauta highway
(Peru)
A. eunotus

   A. cf. eunotus (Shahuaya)
   - 

A 73
M. Río Ucayali (Peru)

Río Shahuaya (near Pucallpa, Peru)
A. cf. eunotus
   (Orangebauch/Orange-belly)
A 76 L. Río Ucayali
(between Nauta & Iquitos, Peru)
A. cf. eunotus
   (Orangeschwanz/Orange-tail)







   A. cf. eunotus (Santa Ana)
A. sp. "Willy" (in trade,
not same as Aqualog);
A. sp. "Orange";
A. sp. "Red & Brown";
A. sp. "Rummy Nose";
A. sp. "Red cruzi";
A. sp. "Paracas";
A. sp. "Jumbo"
A 75








A 74
L. Río Ucayali (Peru)








L. Río Ucayali (near Santa Ana, Peru)
A. cf. eunotus (Río Huallaga)    -  Río Huallaga (near Tarapoto, Peru)
A. cf. eunotus
   (Schwanzstreifen/Tail-stripes)

   A cf. eunotus (Huanta)
A. sp. aff. eunotus (Koslowski);
A. sp. "Albertini"
A 77


   -
L. Ucayali (Peru)?


L. Río Orosa near Huanta (Peru)
A. cf. eunotus (Shishita)    - M. Río Shishita (Peru)
A. sp. "Alto Tapiche" A. sp. "Río Tapiche"    - U. Río Tapiche (Peru)
A. sp. "D16"    - R. Jutai (Brazil)
A. sp. "Nadelstreifen/Needle-stripes" A 78 Río Amazonas (near Leticia, Colombia)
A. sp. "Spaltenfleck/Split-spot" = A. cf. eunotus (Orangeschwanz)?    -  Unknown; exported from Peru
A. sp. "Tahuayo" A. eunotus (Chang, 1996);
(Römer, 1998 & 2000 in part)
A 79 Río Tahuayo (L. Ucayali, Peru)


A.-regani-Group

A.-eunotus-Complex

A.-cruzi-Subcomplex

    Name     Other Names A-No.     Distribution
A. aguarico    -  Río Lagartococha, M. Río Aquarico,
(Ecuador)
A. cf. aguarico (Cuyabeno Reserve)    -  Río Aquarico, Ecuador
(west of Pantoja, Peru)
A. cf. aguarico (Güeppí Reserve) A. sp. "Swordtail-cruzi"    -  Río Aquanegro (north of Pantoja, Peru)
A. cruzi A. sp. "Parallelstreifen/
Parallel-striped"
A 83 Río Mazán (L. Río Napo, Peru)
A. cf. cruzi (San Rafael) A. sp. "Triangle Caudal Spot"    - L. Río Cururay near San Rafael (Peru)
A. cf. cruzi (Vaca Cocha)    -  M. Río Napo
(near Nueva Bella Vista, Peru)
A. sp. aff. cruzi (Picuruyacu) = A. sp. "Nanay"??    -  Lakes beside the Río Picurayacu
(Iquitos, Peru)
A. playayacu

   = A. sp. "Blutauge/Blood-eye"?

   = A. sp. "Caqueta"?
   - 

   - 

A 84
U. Río Napo (Ecuador & Peru)

R. Playayacu (Ecuador/Peru border)

Río Caquetá (Colombia)
A. sp. "D2"

   A. sp. "D2b"
   - 

   - 
R. Caquetá (Colombia)

R. Caquetá (Colombia)
A. sp. "D5"    -  R. Caquetá (Colombia)
A. sp. "D8"    -  R. Caquetá (Colombia)
A. sp. "D15"    -  U. R. Putumayo;
(Puerto Leguízamo, Colombia)
A. sp. "Galaxis" = A. sp. "Nanay" (Melgar)?;
= A. sp. "Putumayo"?
   -  Río Amazonas near Iquitos (Peru)
(Commercially used for several species)
A. sp. "Marandu"    -  Exported from Peru
A. sp. "Nanay" (Melgar)




   = A. sp. "Orangestreifen/
      Orange-stripes"?
A. sp. "Melgar";
A. sp. "Bleeding/Bleeding-heart";
A. sp. "Makurde";
A. sp. "Willy" (Aqualog)
A 82




A 81
U. Río Nanay (Peru)




Río Nanay? (Peru)
A. cf. sp. "Nanay" (Quebrada Yanayacu)    -  L. Quebrada Yanayacu
(L. Río Curaray, Peru)
A. sp. "Napo 1"    -  L. Río Napo (Peru)
A. sp. "Napo 3"    -  L. Río Napo (Peru)
A. sp. "Putumayo" (Algodon II)


   A. sp. "Putumayo" (Río Ampiyacu)
A. sp. "Algodon II";
A. sp. "San Juan"

A. sp. "Parallel-stripes" in Linke & Staeck
A 85


A 86
Río Algodon (Río Putumayo, Peru)


Río Ampiyacu (Peru)
A. cf. sp. "Putumayo" (Red/Blue Face)    -  U. Río Napo, Ecuador/Peru border west of Pantoja (Peru)


A.-regani-Group

A.-eunotus-Complex

A.-ortegai-Subcomplex

    Name     Other Names A-No.     Distribution
A. amoena = A. cf. ortegai (Pebas)?    - Río Ampiyacu near Pebas (Peru)
A. ortegai









   = A. cf. ortegai (w/caudal spot)?
A. sp. "Papagei/Parrot";
A. sp. "Algodon I";
A. sp. "Galaxis";
A. sp. "Algunas";
A. sp. "Nanay" (Römer);
A. sp. "Papagallo";
A. sp. "Frank";
A. sp. "Larinus II";
A. sp. "Larrini"

A. sp. "Papagei/Parrot"
(w/caudal spot)
A 91









A 92
Quebrada Paucaryacu,
M. Río Ampiyacu (Peru)








Unknown (Peru)
A. cf. ortegai (Emerald Pebas) A. sp. "Emerald"    -  Not disclosed; river between Orán
and Pebas (Peru)
A. cf. ortegai (Pebas)




   A. cf. ortegai (Morado/Purple)


   A. cf. ortegai (Naronha)

   A. cf. ortegai Pebas
      (w/caudal spot)
A. sp. "Pebas";
A sp. "Ampiyacu";
A. sp. "Dolly";
A. sp. "Big-foot"

A. sp. "Pebas" (Morado/Purple);
A. sp. "Morado"

A. sp. "Pebas" (Naronha)

A. sp. "Pebas" (w/caudal spot)
A 87




A 89


A 88

A 90
Río Ampiyacu (Pebas, Peru)




Río Ampiyacu (Pebas, Peru)


Río Ampiyacu (Pebas, Peru)

Río Ampiyacu (Pebas, Peru)
A. cf. ortegai (Roca Eterna) A. sp. "Roca Eterna"    -  Not disclosed; river between
Orán and Pebas (Peru)
A. cf. ortegai (Sumon) A. sp. "Montenegro";
= A. ortegai (w/caudal spot)?
A 92? M. Río Yahuasyacu/L. Río Sumon (near Brilla Nueva, Peru)
A. sp. "Totoya"    - Río Algodon (Peru)


A.-alacrina-Group


    Name     Other Names A-No.     Distribution
A. alacrina    -  Río Ariari (Río Guaviare, Colombia)
A. cf. alacrina
   (Ortegueza/Paratype Form)
A. sp. "Rotpunkt" (Ortugueza);
A. sp. "Montañita";
A. sp. "Red-point I";
A. sp. "Caquetá 1"
A 107 Río Ortegueza (Río Caquetá, Colombia)
A. cf. alacrina (Guayas) = A. alacrina?;
A. sp. "Rotpunkt" (Guayas);
A. sp. "Florencia";
A. sp. "Red-point II";
A. sp. "Caquetá 2";
A. sp. "D18"
A 108 Río Guayas (U. Río Caquetá, Colombia)
A. cf. alacrina (Rotpunkt) A. sp. "Schwarzsaum/
Black-rimmed";
A. sp. "Puerto Nariño";
A. sp. "Blue-cheek";
"A. weisei"
A 106 U. Río Caquetá (Colombia)
A. cf. alacrina (Tena)    -  Tributaries of the Río Napo near Tena (Ecuador)
A. sp. "D1"    -  R. Caquetá (Colombia)
A. sp. "D4"    -  R. Caquetá (Colombia)
A. sp. "Kiemenfleck/Gill-spot"
                                  (Inirida)

   A. sp. "Kiemenfleck/Gill-spot
                                  (Vaupes 1)

   A. sp. "Kiemenfleck/Gill-spot
                                  (Vaupes 2)
A. sp. "D37"    - 


   -


   -
U. R. Inirida (Colombia)


U. R. Vaupés (Colombia)


U. R. Vaupés (Colombia)
A. sp. "Mitú" A. sp. "Río Vaupés";
A. sp. "D6";
A. sp. "Slender Gold"
A 109 Río Vaupés (Mitú, Colombia)
A. sp. "Ouaou"    - Unknown location in Colombia


A.-macmasteri-Group

A.-hongsloi-Complex

    Name     Other Names A-No.     Distribution
A. helkeri    - L. Río Sipapo (L. Río Cuao, Venezuela)
A. hongsloi (Rotstrich/Red-streak)


   A. cf. hongsloi (Grau/Gray)
A. sp. "Rotstrich/Red-streak"


Similar to life colors of the holotype
A 110


   -
L. Río Vichada, Guaviare, & Meta
(Colombia & Venezuela)

Colombia
A. cf. hongsloi (Capanaparo) A 111 Río Capanaparo;
Río Suapore, (Venezuela)?
A. cf. hongsloi (Cataniapo)


   A. cf. hongsloi (Maripa)
A. hongsloi (Río Cataniapo) A 113


A 114
Río Cataniapo (Venezuela)
(east of Porto Ayacucho, Colombia)

L. Río Caura (near Maripa);
Near Las Guabinas (Venezuela)?
A. cf. hongsloi (Gold-orange) A. hongsloi
(Orangegelb/Orange-yellow);
A. pleurotaenia (mis-ID)
A 115 Río Claro?; Cinaruco? (Venezuela)
A. cf. hongsloi (Río Claro) A 112 Río Claro (Venezuela)
A. sp. "Rautenband/Diamond-band" A. cf. helkeri;
A. sp. aff. "Mitú"
   - Río Vaupés; Río Tiquié (Colombia)


A.-macmasteri-Group

A.-macmasteri-Complex

A.-macmasteri-Subcomplex

    Name     Other Names A-No.     Distribution
A. guttata


   A. sp. "Hochflossen/High-finned"



A. reitzigi (mis-ID)
A 126


   -
Río Morichal Largo (Orinoco delta, Venezuela)

Unknown (Venezuela)?
A. macmasteri (wild yellow)


   A. macmasteri
      (Rotrücken/Red Shouldered Mac)
A. sp. "Giant Yellow";
A. sp. "Red & Yellow"

A. ornatipinnis (mis-ID);
A. sp. "Red & Blue";
A. sp. "Canoga Park";
A. sp. "Flame";
A. sp. "Meteor"
A 120


A 121
U. Río Meta (Colombia)


U. Río Meta (Colombia)
A. cf. macmasteri/viejita (wild) wild hybrid? A 122 imported from Colombia
A. viejita A. sp. "Rotsaum/Red-seam";
A. viejita Color Form I
A 123 U. Río Meta (Puerto Gaitán, Colombia)
A. sp. "D21"    -  R. Vichada near Puerto Carreño (Colombia)
A. sp. "D23"    -  R. Guarrojo? (Colombia)
A. sp. "D24" = Colorful morph of D23?    -  R. Guarrojo? (Colombia)
A. sp. "D25"    -  Colombia
A. sp. "D26"    -  Colombia
A. sp. "D27"    -  Colombia
A. sp. "D28"    -  Colombia
A. sp. "D29" = A. sp. "Schwarzkehl"?    -  Colombia
A. sp. "D30"    -  Colombia
A. sp. "D31"    -  Colombia
A. sp. "D32"    -  Colombia
A. sp. "D33"    -  Colombia
A. sp. "D34"    -  Colombia
A. sp. "Nana"    -  Colombia?
A. sp. "Rotflecken/Red-flecked" A. viejita Color Form II;
A. sp. "Schwarzkehl/Rotflecken"
A 125 Headwaters between R. Muco & R. Guarrojo (U. Río Vichada, Colombia)
A. sp. "Schuppenfleck/Scale-spot"    -  L. Orinoco, Colombia? Venezuela? (possible hybrid)
A. sp. "Schwarzkehl/Black-throat" A. viejita Color Form III A 124 U. Rio Manacacias, (U. Rio Meta, Colombia)
A. sp. "Zickzack/Zigzag" A. sp. "Kolumbien" A 127 Colombia


A.-macmasteri-Group

A.-macmasteri-Complex

A.-hoignei-Subcomplex

    Name     Other Names A-No.     Distribution
A. caudomaculata    -  L. Río Orinoco (Ríos Candela, Arature, Upata and Caura, Venezuela)
A. hoignei

   A. hoignei
      (Broad Red Caudal Stripes)

   A. hoignei (Tinaquillo)
A 116

A 118


A 116
L. - M. Río Orinoco (Venezuela)

Río Portuguesa Venezuela?


Near Tinaquillo (Venezuela)
A. cf. hoignei (Río Arauca) A. sp. "Tamé"

= A. sp. "D13"?;
= A. sp. "D17"?;
= A. sp. "Casanare"?
A 117

   -
Río Tamé (U. Río Arauca, Colombia)

R. Casanare, near Tamé (Colombia)
A. intermedia    -  L. Río Orinoco between Río Caura & delta (Venezuela)
A. nororientalis A. hoignei (Uracoa)    -  Gulf of Paria (Río San Juan) and lower
Río Orinoco; Río Uracoa (Venezuela)
A. pedunculata A. sp. "Río Caura";
= A. sp. "D7"?
A 119 L. Río Orinoco (U. & M. Río Cataniapo &
Río Caura, Venezuela); E. Colombia
A. sp. "D7" = A. pedunculata?    - R. Vichada, Cumaribo (Colombia)
A. sp. "D12"





    A. cf. sp. "D12"


    A. sp. "D36"


    A. cf. sp. "Pacman" (Japan)
A. sp. "Guainia";
A. sp. "Pacman"
A. sp. "Guaviare"
   - 





   -


   -


   -
Río Vichada drainage (Colombia)?
Northern tributaries to
the Río Guaviare?
Southern tributaries to
the Río Guaviare?

Northern tributaries to
the Río Guaviare

Southern tributaries to
the Río Guaviare

Río Guaviare drainage (Colombia)
Imported to Japan in 2017
A. sp. "D47"    -  Colombia


A.-macmasteri-Group

A.-macmasteri-Complex

Affiliation Uncertain

    Name     Other Names A-No.     Distribution
A. sp. "Casanare" A. cf. guttata "Casanare";
A. sp. "Macguttata";
= A. cf. hoignei (Arauca)?;
= A. sp. "D13"?;
= A. sp. "D17"?
   - R. Casanare (R. Meta system, Colombia)
A. sp. "D13" = A. cf. hoignei (Arauca)?;
= A. sp. "D17"?;
= A. sp. "Casanare"?
   - R. Casanare, near Tamé (Colombia)
A. sp. "D17" = A. cf. hoignei (Arauca)?;
= A. sp. "D13"?;
= A. sp. "Casanare"?
   - R. Casanare, near Arauca, Colombia
A. sp. "D19" = A. sp. "D12"?    - R. Vichada drainage (Colombia)
A. sp. "D35"    - Colombia


A.-pertensis-Lineage


Apistogramma wapisana

    Name     Other Names A-No.     Distribution
A. wapisana A. sp. "Balzfleck/Courting-patch" A 140 Rio Uraricuera (U. Rio Branco), near Barcelos (M. Rio Negro, Brazil)


A.-steindachneri-Group


    Name     Other Names A-No.     Distribution
A. hippolytae


   A. cf. hippolytae (Rio Tefé)
A. sp. "Zweipunkt/Two-spot" A 134


A 135
M. & L. Rio Negro;
Rio Manacapuru (Brazil)

Rio Tefé (Brazil)
A. cf. hippolytae (Rio Trombetas) A 133 M. Rio Trombetas (Brazil)
A. rupununi


   A. cf. rupununi (Boa Vista)

   A. cf. rupununi (Rio Anauá)
A. "ortmanni rupununi";
A. sp. "Zweifleck/Two-patch"
A 130


A 131

A 132
Guyana;
Rio Branco (Brazil)

U. Rio Branco (Brazil)

Rio Anauá (M. Rio Branco, Brazil)
A. steindachneri A. ornatipinnis;
A. wickleri;
A. ortmanni (mis-identification)
A 138 Guyana;
Surinam;
E. Venezuela
A. cf. steindachneri (Río Cuyuni) A 139 Río Cuyuni (Venezuela)
A. sp. "Rio Preto" (do Candeias) A. sp. "Rio Preto de Candeias" A 136 Rio Preto do Candeias (U. Rio Madeira, Brazil)
A. sp. "rupununi-Savanne" A 137 U. Essequibo River (Guyana);
U. Rio Branco (Brazil)


A.-pertensis-Group

A.-pertensis-Complex

    Name     Other Names A-No.     Distribution
A. angayuara A. sp. "Trombetas 1";
= A. sp. "Kupferrücken/
Copper-backed"?
A 145 M. Rio Trombetas near Porteiras (Brazil)
A. inornata


   A. cf. inornata (Guariquito)
A. sp. "Maniapuré";
A. sp. "Piacoa"

A. sp. "Guariquito"
A 142


A 143
Río Maniapuré;
Río Tiquiré; Piacoa (Venezuela)

Río Guariquito (M. Río Orinoco, Venezuela)
A. meinkeni

   A. sp. aff meinkeni
A 144 M. Rio Uaupés (Brazil)

M. Rio Negro (Brazil)?
A. minima



   A. sp. "D20"
A. sp. "Weißsaum/White-seam";
A. sp. "Minima";
A. sp. "Tavi"
A 141



   - 
U. - M. Orinoco between Pto. Ayacucho
& San Fernando de Atabapo (Venezuela
& Colombia)

R. Vichada, Puerto Carreño (Colombia)
A. pertensis A. sp. "Rusty" A 146 U? - L. Rio Negro;
M. Amazon? (Brazil)
A. cf. pertensis




   A. cf. pertensis (Arapiuns)

   A. cf. pertensis (Japurá)

   A. cf. pertensis (Lower Negro)

   A. cf. pertensis (Pimental)

   A. cf. pertensis (Preto do Unini)

   A. cf. pertensis
      (sp. aff. meinkeni)

   A. cf. pertensis (Tefé)

   A. cf. pertensis (Tromebtas)
A. meinkeni (mis-ID);
A. cf. meinkeni





A. pertensis (mis-ID)

A. pertensis (mis-ID)

A. sp. "Pimental"

A. pertensis (Römer)

A. sp. aff. meinkeni
(Römer CA2)
   -




A 149

   -

A 147

A 150

   -

   -


A 148

A 152
Tributaries of the Solimões/Amazon
between the Rio Japurá
and Rio Tapajós (Brazil)


L. Rio Arapiuns (Brazil)

L. Rio Japurá (Brazil)

L. Rio Negro (Brazil)

Rio Pimental (L. Rio Tapajós, Brazil)

Rio Preto do Unini (M. Rio Negro, Brazil)

M. Rio Negro? (Brazil)


L. Rio Tefé (Brazil)

L. Rio Trombetas (Brazil)
A. cf. pertensis
   (Orangesaum/Orange-rimmed)
A. sp. "Orangesaum/
Orange-rimmed"
A 151 Río Ventuari (Guachapana, Venezuela)
A. sp. "Erdfresser/Earth-eater"



   A. cf. sp. "Erdfresser/Earth-eater"
      (w/lateral spot)
A. sp. "Lyretail Purus";
A. sp. "Rondonia";
A. pulchra (mis-ID);
A. hoignei (Uracoa)
A 154



A 155
U.-L. Rio Purus;
Lago Manacapuru (Brazil)


U.-L. Rio Purus (Brazil)
A. sp. "Schwanzzipfel/Tail-point" A. cf. pertensis (Upper Negro Lancetail) A 153 U. Rio Negro (Brazil)


A.-pertensis-Group

A.-velifera-Complex

    Name     Other Names A-No.     Distribution
A. velifera A. sp. "Vierstreifen/Four-stripes";
= A. sp. "Ventuari 1"?
A 163 M. Río Orinoco (Venezuela) to M. Uaupés? (Brazil)
A. sp. "D40"    - Río Guaviare (Colombia)
A. sp. "Putzer/Cleaner" A. sp. "São Gabriel Lyretail";
A. sp. "San Gabriel"
A 162 U. Rio Negro (São Gabriel da Cachoeira, Brazil)


A.-iniridae-Group


    Name     Other Names A-No.     Distribution
A. flabellicauda A. uaupesi (Bork)



= A. sp. "D42"?;
= A. sp. "D44"?
A 156
(Bork
Photo)

   -
U. Río Orinoco between Río Ventuari
& Río Autana (Venezuela);
Rio Uaupés (Brazil)

U. Río Negro (Colombia)
A. iniridae

   A. cf. iniridae
      (w/red gill cover markings)
A. sp. "Fadenflossen/Thread-fin" A 159

A 160
L. Río Inirida (L. Río Guaviare, Colombia)

U. Orinoco (Colombia)
A. lineata    -  Río Atabapo from mouth to Río Atacavi (Colombia & Venezuela)
A. uaupesi A. sp. "Rotkeil/Red-wedge";
A. sp. "Arica";
A. sp. "Ararira";
A. sp. "Felsen/Rock";

A. sp. "Ventuari 2"
= A. flabellicauda?;

= A. sp. "D42"?;
= A. sp. "D44"?
A 156 Río Uaupés/Vaupes (Brazil/Colombia)




L. & M. Río Ventuari (Venezuela)


U. Río Negro (Colombia)
A. sp. "Blutkehl/Cutthroat" A 157 Río Atabapo;
L. Río Inirida;
L. Río Guaviare (Colombia)
A. sp "D42" = A. uaupesi?    - U. Río Negro (Colombia)
A. sp "D44" = A. uaupesi?    - U. Río Negro (Colombia)
A. sp "D51"    - Eastern Colombia
A. sp. "Içana" A 161 Rio Içana (U. Rio Negro, Brazil)
A. sp. "Río Casiquiare" = A. sp. "D42"?    - Río Casiquiare (Venezuela)
A. sp. "Segelflossen/Sail-fin"

   A. sp. "Autana"
A 158

   -
M. Río Orinoco (Venezuela)

R. Autana (Venezuela)


A.-trifasciata-Lineage

A.-trifasciata-Sublineage

Apistogramma arua

    Name     Other Names A-No.     Distribution
A. arua A. sp. "Aruã" A 170 Rio Arapiuns (L. Rio Tapajós, Brazil)


A.-brevis-Group


    Name     Other Names A-No.     Distribution
A. brevis A 168 Rio Uaupés (Brazil)
A. megaptera A. cf. "Breitbinden (Kurzlappen)/
Short-lappets";
A. brevis (mis-ID)
A 166 Ríos Inirida and Mavaca (Colombia);
Sipapo System (Venezuela);
U. Rio Negro?(Exported from São Gabriel da Cachoeira, Brazil)
A. personata


   A. cf. personata (Colombia)



A. personata Mitú
A 171


A 172
M. & U. Río Uaupés/Vaupés
(Brazil/Colombia)

Río Vaupés (Mitú, Colombia)
A. piaroa



   A. cf. sp. "Breitbinden" (Staeck)
A. sp. "Breitbinden/Broad-banded";
A. sp. "Caño Morrocoy"
A 164



A 167
U. Rio Negro (Brazil);
U. Río Orinoco (Colombia, Venezuela)


Imported from Colombia
A. cf. piaroa (Rio Negro) A. sp. "Windisch";
A. sp. "Breitbinden" (São Gabriel);
A. sp. "Schwanzfleck/Tail-spot"
A 165 U. Rio Negro near São Gabriel da Cachoeira (Brazil);
U. Río Negro (Colombia)
A. cf. piaroa (Yellow Breitbinden) A. sp. "D11"    -  R. Vichada, Cumaribo (Colombia)
A. sp. "Alto Vaupes I" A. sp. "D39"    -  U. R. Vaupés (Colombia)
A. sp. "Tiquié" A. sp. "Tiquié 1" A 169 Rio Tiquié (M. Rio Uaupes, Brazil)
A. sp. "Yellow-chin" A. cf. personata (Rio Içana)    -  Río Içana (Brazil, Colombia)?


A.-brevis-? / A.-cacatuoides-Group?


    Name     Other Names A-No.     Distribution
A. salpinction


   A. cf. salpinction (Itapeaçu)
A. sp. "Upper Trombetas";
A. sp. "Trombetas 2"
A 173


   - 
M. Rio Trombetas (Brazil)


Mouth of Rio Madeira
near Itapeaçu (Brazil)
A. sp. "Doppelfleck/Double-spot" A 192 Rio Iriri? Rio Negro (Brazil)


A.-sp.-"D10"-Group


    Name     Other Names A-No.     Distribution
A. sp. "Alto Vaupes II" A. sp. "D50";
A. sp. "Unibrow"
   - Río Vaupés drainage (Colombia)
A. sp. "D10"    - Río Caqueta drainage (Colombia)
A. sp. "Jirijirimo"    - Jirijirimo waterfall, M. Río Apaporis (Colombia)


A.-nijsseni-Group


    Name     Other Names A-No.     Distribution
A. baenschi A. sp. "Inka/Inca";
A. sp. "Inka 50";
A. sp. "Hifin nijsseni"(1)
A 188 M. Río Huallaga
(between Tarapoto & Yurimaguas, Peru)
A. feconat A. sp. "Zwilling/Twin";
A. sp. "Hifin nijsseni" (2);
A. cf. payaminonis (Melgar)
A 181 U. & M Río Tigre (Peru)
A. martini A. sp. "Leierschwanz";
A. sp. "Gabelschwanz II – Blue"
A 187 U. Río Nanay (Peru)
A. nijsseni   A 180 Small quebradas east of Jenaro Herrera,
Río Ucayali drainage, Peru
A. panduro


   A. cf. panduro
      (striped tail/long caudal spot)

   A. cf. panduro
      (striped tail/red dorsal blotch)
A. sp. "Pandurini";
A. sp. "Blue-sky"
A 183


A 184


A 185
Río Tahuayo (L. Río Ucayali, Peru)


Río Tamshiyacu (L. Río Ucayali)


Río Tamshiyacu (L. Río Ucayali)?
A. pantalone A. sp. "Lyretail panduro";
A. sp. "Leierschwanz";
A. sp. "Gabelschwanz";
"Color Form I – Yellow, Peru (3)"
A 186 U. Río Nanay (Peru)
A. payaminonis   A 179 U. Río Napo (Ecuador);
U. Río Aguarico (Equador/Colombia)
A. rositae A. sp. "Harlekin/Harlequin";
A. sp. "Urias";
A. sp. "Hifin panduro";
A. sp. "Black-triangle"
A 189 L. Río Marañon & mouth of Río Huallaga (Peru)
A. wolli A. cf. payaminonis (Pantoja);
A. cf. payaminonis (Kästner);
A. cf. payaminonis (Peru)
A 178 U. Río Napo near Pantoja,
Peru/Ecuador border
A. sp. "D3" A. cf. payaminonis (Colombia)    - Río Caqueta (Colombia)
A. sp. "Matses" A. sp. "nijsseni-New"
(no tail pattern)
   - Río Galvez (Peru)
A. sp. "Oregon" A. sp. "Jav" A 182 M. Río Itaya (Peru)


A.-cacatuoides-Group


    Name     Other Names A-No.     Distribution
A. allpahuayo A. sp. "Schwarzkinn/Black-chin";
A. juruensis (mis-ID);
A. sp. aff. juruensis;
A. sp. "Pucallpa";
A. sp. "Fanta"
A 198
(Blue)


A 199
(Yellow)
M. Río Ucayali Pucallpa? (Peru)
A. cacatuoides



   A. cf. cacatuoides (Rio Careiro)

   A. cf. cacatuoides
      (Rio Manacapuru)

   A. cf. cacatuoides (Río Napo)


   A. cf. cacatuoides
      (Río Putumayo)

   A. cf. cacatuoides (Rio Juruá)

   A. cf. cacatuoides (Solano)
"Großmaul Apisto";
"Cockatoo Dwarf Cichlid";
A. borellii (mis-ID)









A. cf. cacatuoides (Estrecho)
A 200



   -

   -


   -


A 202


A 201

   -
Río Ucayali & Amazonas (Peru);
Rio Solimões (Brazil)


Rio Careiro (Brazil)

Rio Manacapuru (Brazil)


Río Napo between Río Curaray and Río Aquarico (Peru)

Río Putumayo (Peru/Colombia/Brazil?);
Río Algodon (Peru)

Rio Juruá (Brazil)

M. Río Caqueta (Colombia)
A. cf. cacatuoides (Aguaytia)    - Río Aguaytia (U. Río Ucayali, Peru)
A. cf. cacatuoides (Pucallpa) = A. sp. "Superorbital"?    - M. Ucayali near Pucallpa (Peru)
A. juruensis A 197 U. Rio Juruá (Cruzeiro do Sul, Brazil)
A. cf. juruensis (Peru) = A. sp. "Superorbital"?    - Río Purus; Río Javari (Peru/Brazil)?
A. luelingi


   A. cf. luelingi
      (Puerto Maldonado)
A 195


A 196
Río Madre de Dios (Peru/Bolivia);
U. Río Mamoré (Bolivia)

Puerto Maldonado (Peru)
A. cf. luelingi (Cristal) A. sp. "Cristal"    - Río Madre de Dio near Puerto Maldonado (Peru)
A. sororcula A. cf. staecki (Guaporé);
A. sp. "Río Guaporé";
A. sp. "Guaporé 2"
A 193 U. & L. Río Guaporé (Bolivia & Brazil)
A. staecki A. sp. "Querstreifen/
Vertical-striped"
A 194 L. Río Mamoré (Bolivia & Brazil)
A. sp. "Bolivia Fire-Fin" = A. rositae?    - Unknown; Río Madre de Dios (Bolivia)?
A. sp. "Cruzeiro" A. sp. "Goias Redtail";
A. sp. "Juruá";
A. sp. "Juruá Emerald"
A 191 Rio Juruá (Pernambuco, Brazil)
A. sp. "Rocafuerte" A. sp. "Shally"    - U. Río Napo between Rocafuerte
(Ecuador) and Pantoja (Peru)
A. sp. "Superorbital" A. cf. cacatuoides (Río Acre);
A. cf. juruensis (Peru);
= A. cf. cacatuoides (Pucallpa)?
   - U. Río Acre (Peru)?;
Río Purus?; Rio Javari? (Peru/Brazil)


A.-norberti-Group


    Name     Other Names A-No.     Distribution
A. norberti A. sp. "Großmaul/Big-mouth" A 190 Río Tamshiyacu, Río Tahuayo
(L. Río Ucayali, Peru)
A. cf. norberti (Lyretail) A. cf. norberti (Leticia);
A. sp. "New 2";
A. sp. "Jutai New 2";
A. sp. "High-fin-norberti"
   - Tributaries of the Colombian Amazon
near Leticia, Colombia


A.-atahualpa-Group

A.-atahualpa-Complex

    Name     Other Names A-No.     Distribution
A. atahualpa



    A. cf. atahualpa (w/tail spots)
A. sp. affin. payaminonis;
A. sp. "Sunset/Sunset-norberti";
A. sp. "Goldbauch/Gold-belly"

A. sp. aff. atahualpa (Melgar)
(w/tail spots)
A 175



A 176
Río Nanay (Llanchama, Peru)



Río Nanay (Peru)
A. huascar



   A. cf. huascar (Aleman)

   A. cf. huascar (Christoffersen)
A. cf. atahualpa
(Schwanzzipfel/ Tail-points);
A. sp. "Sun-pin";
A. sp. "Sunrise"
A 177



   -

   -
Río Nanay (Peru)



Undisclosed; U. Río Tapiche (Peru)

Unknown; probably Río Nanay (Peru)


A.-atahualpa-Group

A.-barlowi-Complex

    Name     Other Names A-No.     Distribution
A. barlowi A. sp. "Maulbrüter/Mouthbrooder";
"Brustband/Breast-band";
"Glaser";
"Red-face";
"Half Orange Tail"
A 174 Río Ampiyacu;
L. Río Juruá? (Peru)
A. sp. aff. barlowi (Oran)    - Undisclosed river east of Oran (Peru)
A. megastoma A. sp. "Kelleri";
A. sp. "Leticia";
A. sp. "Jutai";
A. cf. sp. "Keller" (Jutai);
A. sp. "Diamond-face"
   - Leticia, (Colombia);
R. Jutai (Brazil)?


A.-trifasciata-Group


    Name     Other Names A-No.     Distribution
A. erythrura A. sp. "Mamoré";
A. maciliensis (mis-ID)
A 207 M. & L. Guaporé (Bolivia & Brazil);
L. Mamoré (Brazil)
A. trifasciata




   A. trifasciata maciliensis


   A. cf. trifasciata (Argentina)

   A. cf. trifasciata
      (Northern Pantanal)

   A. cf. trifasciata (Río Guaporé)





A. trifasciata "haroldschultzi";
A. maciliensis
A 204
A 205
A 206


   -


A 205

A 204


A 206
Río Paraguay (Brazil, Paraguay,
& Argentina);
U. & M. Rio Guaporé (Brazil & Bolivia);
Río Beni (Bolivia)

Lakes in the M. Río Guaporé
(Bolivia & Brazil)

L. Río Paraguay (Argentina)

U. Rio Paraguai (Brazil)


U. & M. Río Guaporé
(Brazil & Bolivia)


A.-agassizii-Sublineage

A.-elizabethae-Group


    Name     Other Names A-No.     Distribution
A. elizabethae


   A. cf. elizabethae
A. sp. "Rußkopf/Sooty-head";
= A. sp. "Frank"?

A. elizabethae w/damaged dorsal fin?
A 208


   -
Río Uaupés/Vaupés (Brazil/Colombia);
Rio Içana (Brazil)

Rio Negro (Brazil)?


A.-gibbiceps-Group


    Name     Other Names A-No.     Distribution
A. gibbiceps A 209 M. Rio Negro & Rio Branco (Brazil)
A. roraimae A 210 Río Uraricoera & U. Rio Branco
above Boa Vista (Brazil/Guyana)


A.-bitaeniata-Group

A.-bitaeniata-Complex

    Name     Other Names A-No.     Distribution
A. bitaeniata (Brazilian Color Form)


   A. bitaeniata (Careiro)



   A. bitaeniata (Curuaí)


   A. bitaeniata (Curupira)


   A. bitaeniata ((Juruá))

   A. bitaeniata (Madeira/Autazes)

   A. bitaeniata (Mamori)

   A. bitaeniata (Manacapuru)


   A. bitaeniata (Manaquiri)


   A. bitaeniata (Rio Tefé)
A. klausewitzi;
A. kleei















A. bitaeniata (Mamuri)

A. bitaeniata (Januari);
A. bitaeniata (Manaus)
A 215


   -



   -


   -


   -

   -

   -

A 214


   -


A 213
Rio Tefé to mouth of Rio Tapajós (Brazil)


Right bank (south) tributaries of the
Amazon around Careiro da Várzea (Brazil)

Right bank (south) tributaries of the
Amazon around Curuaí (Brazil)

Igarape Curupira
(L. Rio Cunauaru, Brazil)

L. Rio Juruá (Brazil)

Near Autazes (L. Madeira, Brazil)

Rio Mamori (L. Madeira, Brazil)

Rio Manacapuru (Brazil)


Right bank (south) tributaries of
the Amazon near Manaquiri (Brazil)

Rio Tefé (Brazil)
A. bitaeniata (Peruvian Color Form)

   A. bitaeniata (Curutu)

   A. bitaeniata (Leticia)


   A. bitaeniata (Momón)

   A. bitaeniata (Nanay)

   A. bitaeniata (Nieva)

   A. bitaeniata ((Pastaza))

   A. bitaeniata (Putumayo)

   A. bitaeniata (Requena)


   A. bitaeniata (Shishita)


   A. bitaeniata (Shushupi)

   A. bitaeniata (Río Tigre)




= A. sweglesi?
   -

   -

   -


   -

A 211

   -

   -

   -

   -


   -


   -

   -
Río Amazonas (Peru)

Río Curutu (Río Pavayacu, Peru)

Tributaries of the Amazon
near Leticia (Colombia)

L. Río Momón (Peru)

L. Río Nanay (Peru)

Río Nieva (U. Río Marañón, Peru)

L. Río Pastaza; M. Río Marañón (Peru)

Tributaries of M. Río Putumayo (Peru)

Tributaries of L. Río Ucayali
around Requena (Peru)

M. Río Shishita;
M Río Ampyacu (Peru)

M. Río Ampiyacu (Peru)

L. Río Tigre (Peru)
A. cf. bitaeniata (Yavari) A 212 L. Río Yavari (Brazil/Peru)
A. sp. aff. bitaeniata (Porto Velho) A 216 Rio Madeira?
(Exported from Porto Velho, Brazil)
A. eremnopyge A. sp. "Fresa/Strawberry";
A. sp. "Diamante";
A. sp. "Rotbart/Red-beard";
A. sp. "Tapiche"
A 218 Río Itaya (Peru)
A. sp. "Caño de la Libertad"    - Caño de la Libertad and nearby streams,
Río Apaporis (Colombia)
A. sp. "D9"    - Colombia
A. sp. "D14"    - Colombia
A. sp. "Orangeflossen/Orangefins" A 217 domestic hybrid
bitaeniata x paucisquamis-Complex?


A.-bitaeniata-Group

A.-paucisquamis-Complex

    Name     Other Names A-No.     Distribution
A. mendezi (Cross-striped)



   A. mendezi (Long-stripes)


   A. mendezi (Orange)
 



A. sp. "Langsstreifen/
Long-striped"
A 221



A 220


A 222
Right side tributaries of M. & U.
Rio Negro (Rio Urubaxi to
São Gabriel da Cachoeira, Brazil)

Right side tributaries of M. & U.
Rio Negro (Barcelos to Rio Urubaxi, Brazil)

Rio Negro (Brazil)
A. paucisquamis A. sp. "Glanzbinden/
Shiny-banded"
A 219 M. & L. Rio Negro (Brazil)
A. sp. "Gabelband/Fork-band" A. agassizii
(Gabelband/Fork-band)
   - M. Rio Jaú (Rio Negro System, Brazil)
A. sp. Miuá A. sp. "Rio Uambé";
A. sp. "Rio Mabó";
A . sp. "Cacimbo"
A 223 U. Rio Negro (Igarapé Miuá, Uambé, & Mabóm, São Gabriel, Brazil)
A. cf. sp. Miuá (Demini) A. sp. "Miuá" (Yellow) A 223 Rio Demini (Brazil)


A.-agassizii-Group

A.-pulchra-Complex

    Name     Other Names A-No.     Distribution
A. pulchra A. sp. "Mikschofsky" A 224 U. Rio Madeira (Brazil)
A. cf. pulchra (Aripuanã)

   A. cf. pulchra (Chingarno)
A. sp. "Aripuanã"

A. sp. "Chingarno"
A 225

A 225a
Rio Aripuanã (Rio Madeira, Brazil)

Rio Chingarno (L. Rio Madeira, Brazil)
A. sp. "Abacaxis"


   A. cf. sp. "Abacaxis" (Marimari)
A. sp. "Wilhelmi";
A. cf. pulchra (Abacaxis)

A. cf. pulchra (Marimari);
A. sp. "New Chingarno"
A 227


A 228
M. Rio Abacaxis (L. Rio Madeira, Brazil)


Rio Marimari (L. Rio Madeira, Brazil)
A. sp. "Branco"


   A. cf. sp. "Branco" (Uatumã)
A. cf. pulchra (Branco);
A. sp. "Branco-pulchra"

A. cf. pulchra (Uatumã);
A. sp. "Deep Red"
A 226


   -
Rio Cauamé (U.Rio Branco near
Boa Vista, Brazil)

Rio Uatumã (E. of Manaus, Brazil)


A.-agassizii-Group

A.-agassizii-Complex

    Name     Other Names A-No.     Distribution
A. agassizii


   A. agassizii (Japurá)

   A. agassizii (Juruá)

   A. agassizii (Leticia)


   A. agassizii (Putumayo)

   A. agassizii (Red)


   A. agassizii (Tamshiyacu)

   A. agassizii (Tefé)

   A. agassizii (Tigre)







A. agassizii (Tabatinga)







A. cf. agassizii (Butterfly)



A. agassizii (Flamenco)
A 234


   -

   -

   -


   -

A 234


   -

   -

   -
U. Río Amazonas (Peru) to
L. Rio Solimões (Brazil)

L. Rio Japurá (M. Solomões, Brazil)

L. Rio Juruá (Brazil)

Tributaries of Amazon near
Leticia (Colombia)

M. Río Putumayo (Peru)

Río Nanay, Supay, Ampiyacu,Maniti,
L Negro (Peru & Brazil)

L. Río Tamshiyacu (Peru)

L. Rio Tefé (Brazil)

L. Río Tigre (Peru)
A. cf. agassizii (Ampiyacu) A 241 Río Ampiyacu (Peru)
A. cf. agassizii
   (Broad Black Caudal Seam)

   A. cf. agassizii (Broad Black
      Caudal Seam) – Apeú

   A. cf. agassizii (Broad Black
      Caudal Seam) – Careiro


   A. cf. agassizii (Broad Black
      Caudal Seam) – Curuaí

   A. cf. agassizii (Broad Black
      Caudal Seam) – Itaituba

   A. cf. agassizii (Broad Black
      Caudal Seam) Lagoa Janauacá

   A. cf. agassizii (Broad Black
      Caudal Seam) - Madeira




   A. cf. agassizii (Broad Black
      Caudal Seam) – Manaquiri

   A. cf. agassizii (Broad Black
      Caudal Seam) – Tapajós

   A. cf. agassizii (Broad Black
      Caudal Seam)
      – Tapajós Rotrücken/Red-back

   A. cf. agassizii (Broad Black
      Caudal Seam) – Tocantins


   A. cf. agassizii (Broad Black
      Caudal Seam) – Xingu
= A. parva?


A. agassizii (Rio Apeú)






= A. cf. agassizii (Tapajós)?


= A. cf. agassizii (Tapajós)?





Gold agassizii;
A. cf. agassizii (Porto Velho);
A. agassizii (Nova Olinda);
A. agassizii (Rondonia)


A. cf. gephyra (Manaquiri)


A. agassizii (LakeSantarém);
A. agassizii (Óbidos)





A. agassizii (Belém);
A. agassizii (Tucurui);
A. agassizii Gold-line
   -


   -


   -



   -


   -


   -


A 240





   -


A 236


   -



   -



   -
Right bank (south) tributaries of the
Amazon between Manaus & Belém (Brazil)

Rio Apeú (Brazil)


Right bank (south) tributaries of the
Amazon around Careiro da Várzea
(Brazil)

Right bank (south) tributaries of the Amazon around Curuaí (Brazil)

Rio Tapajós near Itaituba (Brazil)


Small streams entering
Lagoa Janauacá (Brazil)

M. & L. Rio Madeira
(Porto Velho– Manicoré, Brazil)




Right bank (south) tributaries of the
Amazon near Manaquiri (Brazil)

Igarapé Açu, Lago Jacundá
(L. Rio Tapajós, Brazil)

L. Rio Tapajós (Brazil)



L. Rio Tocantins (Brazil)



L. Rio Xingu (Brazil)
A. cf. agassizii (Marañon) A 242 Río Marañon (Peru)
A. cf. agassizii (Netz/Net)


    A. cf. agassizii (Netz/Net)
      - Alenquer

   A. cf. agassizii (Netz/Net)
      – Januari

   A. cf. agassizii (Netz/Net)
      - Jari

   A. cf. agassizii (Netz/Net)
      - Negro

   A. cf. agassizii (Netz/Net)
      - Nhamundá

   A. cf. agassizii (Netz/Net)
      - Trombetas

   A. cf. agassizii (Netz/Net)
      - Urubu



A. agassizii (Bara Mansa)







    
A. agassizii (Caurés);
A. agassizii (Janauacá)
   -


A 235


   -


   -


   -


   -


A 237


   -
Left bank (north) tributaries of the Amazon
between Manaus & Rio Jari (Brazil)

Around Alenquer (Brazil)


L. Rio Januari (Brazil)


L. Rio Jari (Brazil)


Rio Negro below mouth of Rio Branco
& Rio Caurés (Brazil)

L. Rio Nhamundá (Brazil)


L. Rio Trombetas (Brazil)


L. Rio Urubu (Brazil)
A. cf. agassizii (Pastel) A 239 L. Ucayali near Requena & Jenaro Herrera,
Río Tamshiyacu; Río Tapiche? (Peru)
A. cf. agassizii (Purus) A 238 U. Rio Purus (Brazil)
A. gephyra A 231 M. & L. Rio Negro & M. Amazon (Brazil)
A. cf. gephyra (Rio Curuá) A 232 Rio Curuá (Brazil)
A. cf. gephyra (Santarém) A. agassizii Santarem
S03336-4 (SAC II)
A 233 Lago Jurucuí? (Near Santarém, Brazil)?
A. sp. "Blauspiegel/Blue-spangle" A. sp. "Red-fin Rio Xingu";
A. cf. pulchra (Xingu)
A 230 Rio Xingu (near Altamira, Brazil)
A. sp. "Tefé" A. cf. agassizii (Tefé) A 243 Rio Tefé & Lago Catuá (Brazil)
A. sp. "Vitória do Xingu" A. sp. "Xingu IV" A 229 Rio Xingu (near Vitória do Xingu, Brazil)


A.-diplotaenia-Lineage

A.-diplotaenia-Group

    Name     Other Names A-No.     Distribution
A. diplotaenia A. sp. "Doppelband/Double-band" A 128 U. - L. Rio Negro (Brazil),
M. Río Orinoco (Río Inirida, Colombia)
A. cf. diplotaenia (Alto Orinoco) A 129 Río Atabapo (Venezuela)


Affiliation Uncertain

    Name     Other Names A-No.     Distribution
A. parva = A. cf. agassizii
(Broad Black Caudal Seam)?
(type specimens too small to be certain)
   - Rio Capim (Brazil)
A. sp. "D52"    - Colombia


Taeniacara

    Name     Other Names A-No.     Distribution
Taeniacara candidi Apistogramma weisei    - Amazon, Rio Tefé to Rio Tapajós
& L. Rio Negro (Brazil)



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